Of a twinkling light, to cease to twinkle.

(idiomatic) Immediately; instantaneously. / 2007 April 28, Hugh Muir, The Guardian:

Alternative form of in the twinkling of a bedstaff

(archaic) In no time at all; immediately.

(idiomatic, colloquial, humorous) A notional look of anticipation or hope (either for sex or a child) in one's father's eyes at or around the time of one's conception.

Of a person, not yet conceived, particularly when referring to a time period preceding their conception.

One who, or that which, twinkles or winks. / (colloquial) An eye.

plural of twinkler

Alternative form of twinkle toes

(archaic) Third-person singular simple present indicative form of twinkle