wrinkling

The character of these wrinklings, when seen on the flat of the beds, forbids the idea that they are due to end on compression […]

Would she be able to appreciate the subtlety of their advances, if they made any; those little twitches and twists of the lips, those narrowings and rollings of the eye, those wrinklings of the speculative temple, that shrugging of an eye-brow at play?