comma

No points were used by the ancient printers, excepting the colon and the period; but, after some time, a short oblique stroke, called a virgil, was introduced, which answered to the modern comma. In the fifteenth century this punctuation was improved by the famous Aldus Manutius with the typographical art in general; when he gave a better shape to the comma, added the semicolon, and assigned to the former points more proper places.

Commas (Polygonia comma) and Question Marks (Polygonia interrogationis) occur from the Gulf Coast to Canada and west to the Rockies. … Question Marks and Commas are handsome butterflies with burnt orange and black markings. … On the underside of each hind wing of the Comma is a small, distinctive silver hook that resembles a comma.