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Released within the mucus, we discovered three types of undeployed nematocysts, as well as microscopic, motile, cellular masses composed of nematocytes that we formally call cassiosomes. While cassiosomes bear some resemblance to another cnidarian structure originating in mesenteries of the starlet sea anemone Nematostella called nematosomes, the unique traits of cassiosomes in C. xamachana include their release into the water column within mucus, the ability to trap and kill prey as mobile grenades outside of the medusa, their organization as an outer epithelial layer surrounding a mostly empty core (rather than a solid ball of cells), and the presence of centrally-located endosymbiotic Symbiodinium dinoflagellates.
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